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No, I don’t have markup installed (it is there in apps but not installed from the mirror).
I think in my case the screenshot functionality is built into AOSP and the editor you get when tapping the resulting preview overlay in the lower left corner of the screen is part of the “gallery” app since using the “edit” feature from gallery launches the same editor. Maybe GrapheneOS just sets that as the default editor, I’m not sure.
AOSP gallery that comes with GrapheneOS. The app info says it’s called
com.android.gallery3d
. There’s some info here in the docs about the relationship between camera, edit functions, and the gallery app.